Dr. Walter Hendelman

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Dr. Walter Hendelman
Emeritus Professor

BSc (Psychology) - McGill University, Montreal, 1958
MD, CM (Medicine) - McGill University, Montreal, 1962
MEd (Education) - OISE/University of Toronto, 2002

613-562-5800 ext. 5406 (Office)
613-562-5636 (Fax)



  • 1962-63: General Rotating Internship - Royal Victoria Hospital, Montreal
  • 1963-64: Junior Assistant Resident - Montreal Children's Hospital, Pediatric Medicine
  • 1964-65: Postdoctoral Research Fellow of the Muscular Dystrophy Association of Canada: Pasadena Foundation for Medical Research, Pasadena, California.
  • 1965-68: Postdoctoral Fellow of the National Institute of Neurological Diseases and Blindness (U.S.A.): Department of Anatomy, College of Physicians and Surgeons, Columbia University, New York.
  • 1968-2010: Assistant, Associate, Full Professor, Department of Anatomy / Department of Cellular & Molecular Medicine, Faculty of Medicine, University of Ottawa
  • 2010- : Professor Emeritus, Faculty of Medicine, University of Ottawa

Special courses / programs

  • 1994-02: Master of Education program at the Ontario Institute for Studies in Education (OISE), Graduate Department of the University of Toronto in the Higher Education Group, specialization in Health Professional Education.
  • 1996: Sabbatical [6 months]: Development of Instructional Software, University of Washington (Seattle)

Society membership

  • Canada Federation of Biological Sciences (Anatomy) [Emeritus]
  • Society for Neuroscience [Emeritus]
  • Alpha Omega Alpha (Medical Honours Society)

FACULTY OF MEDICINE, University of Ottawa

  • 1997-98: Special Task Force on Electronic Communication Faculty of Medicine]
  • 1997-98: Special Task Force on Information Management [Medical Education]
  • 1997-01: Senate Committee on Teaching (University of Ottawa)
  • 2001-02: Institutional Review: Subcommittee on Basic Science Departments
  • 2003-04: Task Force on Medical Ethics and Humanities program
  • 2001-09: Chair, Professionalism Program for medical students, Subcommittee of the Undergraduate Curriculum Committee

Major scientific research publications

  1. Hendelman, W.J. and Booher, J. 1966. Factors involved in the culturing of chick embryo dorsal root ganglia in the Rose chamber. Texas Reports on Biol. and Med., 24: 83-89.

  2. Pomerat, C.M., Hendelman, W.J., Raiborn, C.W., Jr., and Massey, J.F. 1967. Dynamic activities of nervous tissue in vitro.  In: The Neuron, Ed., H. Hyden, Am. Elsevier Publ. Co., New York, pp. 119-178.

  3. Hendelman, W.J. and Bunge, R.P. 1969. Radioautographic studies of choline incorporation into peripheral nerve myelin. J. Cell Biol., 40: 190-208.

  4. Mire, J., Hendelman, W.J., and Bunge, R.P. 1970.  Observations on a transient phase of focal swelling in degenerating unmyelinated nerve fibers. J. Cell Biol., 45: 9-22.

  5. Hendelman, W.J. 1972. A morphologic study of the effects of LSD on neurons in cultures of cerebellum. J. Neuropath. Exp. Neurol., 31: 411-432.

  6. Wojtowicz, J.M., Marshall, K.C. and Hendelman, W.J. 1977. Depression by magnesium ion of neuronal excitability in tissue cultures of central nervous system. Can. J. Physiol. Pharmacol. 55: 367-372.

  7. Marshall, K.C., Hendelman, W.J., Gimbarzevsky, B. and Wojtowicz, J.M. 1977.  Synaptic transmission in organized cultures of cerebellum.  In "Iontophoresis and Transmitter Mechanisms in the Mammalian Central Nervous System: Eds. R.V. Ryall and J.S. Kelly. Elsevier, pp. 221-223.

  8. Hendelman, W.J., Marshall, K.C., Aggerwal, A.S. and Wojtowicz, J.M. 1977.  Organization of pathways in cultures of cerebellum.  In "Cell, Tissue and Organ Cultures in Neurobiology". Eds. S. Fedoroff and L. Hertz. Academic Press, New York, pp. 539-554.

  9. Wojtowicz, J.M., Marshall, K.C. and Hendelman, W.J. 1978. Electrophysiological and pharmacological studies of the inhibitory projection from the cerebellar cortex to the deep cerebellar nuclei in tissue culture. Neuroscience, 3: 607-618.

  10. Hendelman, W.J. and Aggerwal, A.S. 1980. The Purkinje neuron: I. A Golgi study of its development in the mouse and in culture. J. Comp. Neurol. 193: 1063-1079.

  11. Aggerwal, A.S. and Hendelman, W.J. 1980. The Purkinje neuron: II.  Electron microscopic analysis of the mature Purkinje neuron in organotypic culture. J. Comp. Neurol. 193:         1081-1096.

  12. Hendelman, W.J. and Marshall, K.C. 1980. Axonal projection patterns visualized with horseradish peroxidase in organized cultures of cerebellum. Neuroscience 5: 1833-1846.

  13. Marshall, K.C., Wojtowicz, J.M. and Hendelman, W.J. 1980.  Patterns of functional synaptic connections in organized cultures of cerebellum. Neuroscience 5: 1833-1857.

  14. Marshall, K.C., Pun, R.Y.K., Hendelman, W.J. and Nelson, P.G. 1981 A coeruleo-spinal system in culture.  Science 213: 355-357.

  15. Hendelman, W.J., Marshall, K.C., Ferguson, R. and Carrière, S. 1982. Catecholamine neurons of the central nervous system in organotypic culture. Dev. Neurosci. 5: 64-76.

  16. Marshall, K.C. and Hendelman, W.J. Morphophysiological studies of a culture model of the cerebellum.  In: The Cerebellum; New Vistas. Eds.: S. Palay and V. Chan-Palay, Springer-Verlag Publisher. Exp. Brain Res. Suppl. 6: 1982, pp. 69-74.

  17. Hendelman, W.J. Henderson, I. and Ferguson, R. 1983. LSD-induced morphological alterations in Purkinje neurons of the rat. Exp. Neurol. 82: 478-490.

  18. Hendelman, W.J., Jande, S.S. and Lawson, D.E.M. 1984. Calcium-binding protein    immunocytochemistry in organotypic cultures of cerebellum. Brain Research Bulletin 13: 181-184.

  19. Pun, R.Y.K., Marshall, K.C., Hendelman, W.J., Guthrie, P.B. and Nelson, P.G. 1985.  Noradrenergic responses of spinal neurons in locus-coeruleus-spinal cord co-cultures.  J. Neuroscience, 5: 181-191.

  20. Cardoso, E.R., Peterson, E.W. and Hendelman, W.J. 1985. Sub-pial infiltration of blood products following experimental subarachnoid hemorrhage. Acta Neurochirurgica 76: 140-144.

  21. Hendelman, W.J., de Savigny, N. and Marshall, K.C. 1985. Growth and myelination of explant cultures in defined medium. In Vitro.  Cellular and Developmental Biology (Rapid Communication) 21: 129-134.

  22. Hendelman, W.J. A dissection guide for cerebellum-locus ceruleus organotypic cultures. IN: A Dissection and Tissue Culture Manual of the Nervous System. (eds. A. Shahar, J. de Vellis, A. Vernadakis and B. Haber). 1989. Alan Liss, pp. 16-22.

  23. Humphreys, P. Jones, S., Hendelman, W. 1996.  Three-dimensional cultures of fetal mouse cerebral cortex in a collagen matrix.  Journal of Neuroscience Methods. 66:  23-33.

  24. Humphreys, P. and Hendelman, W.  2000.  Analysis of cerebrocortical neuronal migration in three-dimensional fetal mouse cerebral explants: comparison with in vivo.  International Journal of Developmental Neuroscience, 18: 573-584.

Medical education

  1. Hendelman, W.J. 1990. Videotape Series

    • "The Gross Anatomy of the Human Brain" (4 tapes)
    • "The Anatomy of the Interior of the Human Skull" (1 tape)
  2. Hendelman, W.J. and Boss, M. 1986. Reciprocal peer teaching by medical students in the Gross Anatomy Laboratory. J. Med. Ed. 61: 674-680.

  3. Hendelman, W.J. 1991. A perspective on medical education: A return to clinical residency.  The Pharos. 54(4): 2-7.

  4. Hendelman, W., Sundsten, J.W. and Bolles, J.R. 1997.  Computer-Based Learning of the Nervous System: The Motor System.  Society for Neuroscience (Teaching) 23: 285.

  5. Hendelman, W. and Skinner, C.  2001. Syringomyelia: A problem-based on-line learning module of the spinal cord.  Society for Neuroscience (abstract).

  6. Hendelman, W. and Staines, W.  2002. Neuroanatomy laboratory learning using digital images.  Australian Neuroscience Society (abstract).

  7. C. Skinner and W. Hendelman 2002.   On-Line Problem-Based Learning - University of Ottawa.   Slice-of-Life Workshop.


  1. Walter Hendelman, Darin Davidson, Jennifer Ingram, Isabelle Leblanc, Karine Lortie, Chad Lund, Anne-Marie Therrien, Daniel Trottier, and Linda Peterson. 2002.  Professionalism for Undergraduate Students of the Faculty of Medicine, University of Ottawa.  Association of Canadian Medical Colleges (ACMC).

  2. W. Hendelman and A Byszewski, 2007. A National Survey: Medical Professionalism in Canadian Undergraduate Programs.  Professionalism Resource Group, Association of Faculties of Medicine, Canada (AFMC).  Posted on the AFMC website: [accessed Jan. 29, 2008].

  3. Anna Byszewski, Walter Hendelman, Caroline McGuinty and Geneviève Moineau. Wanted: role models - medical students’ perceptions of professionalism. BMC Medical Education. 2012, 12:115 [doi:10.1186/1472-6920-12-115].

  4. Walter Hendelman and Anna Byszewski. Formation of medical student professional identity: categorizing lapses of professionalism, and the learning environment. BMC Medical Education. 2014, 14:139 [doi:10.1186/1472-6920-14-139].

Book publications

  1. Hendelman, W.J. and Morrissey, J.P. 1987. A Student's Atlas of Neuroanatomy.  University of Ottawa Press.

  2. Hendelman, W.J. and Morrissey, J.P. 1988. A Student's Atlas of Neuroanatomy.  (2nd edition). University of Ottawa Press.

  3. Hendelman, W. J. 1994. A Student's Atlas of Neuroanatomy (3rd edition). W.B. Saunders

  4. Hendelman, W. 2000. Atlas of Functional Neuroanatomy. CRC Press.

  5. Hendelman, W. 2006. Atlas of Functional Neuroanatomy. 2nd edition. CRC Press / Taylor & Francis

  6. Hendelman, W., 2009. Atlante di Neuroanatomia funzionale. Casa Editrice Ambrosiana.

  7. Hendelman, W., Humphreys, P., and Skinner, C. 2010 The Integrated Nervous System: a systematic diagnostic approach. CRC Press / Taylor & Francis

  8. Hendelman, W. Atlas de Neuroanatomie Fonctionelle. [translation] 2013. University of Ottawa Press

  9. Hendelman, W. 2016. Atlas of Functional Neuroanatomy. 3rd edition. CRC Press / Taylor & Francis 


The website for the ATLAS of Functional Neuroanatomy — in both English & French — including video demonstrations of brain anatomy (narrated)


Dr. Walter Hendelman, M.D., C.M., is a Canadian, born and raised in Montreal. He did his undergraduate studies at McGill University in science with honors in psychology. As part of his courses in physiological psychology, he assisted in an experimental study of rats with lesions of the hippocampus, which was then a little known area of the brain. Dr. Hendelman proceeded to do his medical studies at McGill. Subsequently, he completed an internship and a year of pediatric medicine, both in Montreal.

Having chosen the brain as his life-long field of study and work, the next decision involved the choice of either clinical neurology or brain research - Dr. Hendelman chose the latter.  Postgraduate studies continued for four years in the United States, in the emerging field of developmental neuroscience, using the ‘new’ techniques of nerve tissue culture and electron microscopy. Dr. Richard Bunge was his research mentor at Columbia University Medical Center in New York City, while his neuroanatomy mentor was Dr. Malcolm Carpenter, author of the well-known textbook of Human Neuroanatomy.

Dr. Hendelman returned to Canada and has made Ottawa his home for his academic career at the University of Ottawa, Faculty of Medicine, in the Department of Anatomy, now the Department of Cellular and Molecular Medicine. He began his teaching in Gross Anatomy and Neuroanatomy and in recent years has focused on the latter. His research continued, with support from Canadian granting agencies, using nerve tissue culture to examine the development of the cerebellum; later on he became involved in studies on the development of the cerebral cortex.  Several investigations were carried out in collaboration with summer and graduate students and with other scientists. He has been a member of various neuroscience and anatomy professional organizations, has attended and presented at their meetings, and has numerous publications on his research findings.

In addition to research and teaching and the usual academic 'duties', Dr. Hendelman was involved with the Faculty and University community, including a committee on research ethics. He has also been very active in curriculum planning and teaching matters in the Faculty. During the 1990s, when digital technology became available, Dr. Hendelman recognized its potential to assist student learning, particularly in the anatomical subjects and helped bring technology into the learning environment of the Faculty. He also organized a teaching symposium for the Canadian Association of Anatomy, Neurobiology and Cell Biology on the use of technology for learning the anatomical sciences.

In 2002, Dr. Hendelman completed a program in medical education and received a Master's degree in Education from the Ontario Institute of Studies in Education, affiliated with the University of Toronto. In the same year, following retirement, he began a new stage of his career, with the responsibility for the development of a Professionalism Program for medical students at the University of Ottawa.

As a student of the brain, Dr. Hendelman has been deeply engaged as a teacher of the subject throughout his career.  Dedicated to assisting those who wish to learn functional neuroanatomy, he produced several teaching videotapes using anatomical specimens and five previous editions of his Atlas of Functional Neuroanatomy (under various titles). As part of this commitment he has collaborated in the creation of two computer-based learning modules, one on the spinal cord based upon the disease syringomyelia and the other on voluntary motor pathways; both contain original graphics to assist in the learning of the challenging and fascinating subject matter, the human brain.

Most recently, in order to keep pace with the use of technology, Dr. Hendelman guided the development of a web site for the second edition of his Atlas with interactive features. In addition, the 2nd edition has been translated into Italian and now into French and the French edition also has a website. Finally, Dr. Hendelman is a co-author of The Integrated Nervous System: a systematic diagnostic approach, published by CRC Press (2010).

In his non-professional life Walter Hendelman is a husband, a father, an active member of the community, a lover of music, sometimes a choir member and now a member of a community band (trumpet), a commuter cyclist, and an avid skier and skater.